*{box-sizing:border-box}
:root{
  --bg:#0b0e0d;--panel:#101514;--panel2:#151b19;--text:#f2f4ef;--muted:#aab3ad;
  --line:#2a332f;--accent:#d7ff66;--max:1180px
}
html{scroll-behavior:smooth}
body{margin:0;background:var(--bg);color:var(--text);font-family:Manrope,Arial,sans-serif;line-height:1.6}
a{color:inherit;text-decoration:none}
img{display:block;width:100%;height:100%;object-fit:cover}
.container{width:min(var(--max),calc(100% - 48px));margin:auto}
.site-header{position:sticky;top:0;z-index:20;background:rgba(11,14,13,.9);backdrop-filter:blur(14px);border-bottom:1px solid rgba(255,255,255,.07)}
.nav-wrap{min-height:76px;display:flex;align-items:center;justify-content:space-between;gap:32px}
.brand{display:flex;flex-direction:column;line-height:1.05}
.brand span{font-weight:700;letter-spacing:-.03em}
.brand small{margin-top:6px;color:var(--muted);font:11px "DM Mono",monospace;text-transform:uppercase;letter-spacing:.08em}
nav{display:flex;align-items:center;gap:26px;font-size:14px}
nav a{color:#cbd1cd}
nav a:hover{color:#fff}
.nav-cta{border:1px solid var(--line);padding:10px 14px;border-radius:999px}
.hero{padding:88px 0 110px;border-bottom:1px solid var(--line)}
.hero-grid{display:grid;grid-template-columns:1.02fr .98fr;gap:72px;align-items:end}
.eyebrow,.section-label,.project-kicker,.nameplate{font:11px "DM Mono",monospace;text-transform:uppercase;letter-spacing:.12em;color:#a9b3ac}
.eyebrow{margin:0 0 24px}
h1,h2,h3,p{margin-top:0}
h1{font-size:clamp(52px,7vw,92px);line-height:.94;letter-spacing:-.065em;margin-bottom:28px}
h1 em,h2 em{font-style:normal;color:var(--accent)}
h1 strong{font-weight:500}
.hero-lede{max-width:590px;color:#c4cbc6;font-size:18px;line-height:1.65}
.hero-actions{display:flex;gap:12px;flex-wrap:wrap;margin-top:32px}
.button{display:inline-flex;align-items:center;justify-content:center;gap:14px;border-radius:999px;padding:13px 18px;font-size:14px;font-weight:700;border:1px solid var(--line);transition:.2s ease}
.button-primary{background:var(--accent);border-color:var(--accent);color:#10140d}
.button-primary:hover{transform:translateY(-2px);filter:brightness(1.04)}
.button-ghost:hover{border-color:#5b665f}
.hero-meta{display:flex;gap:14px;margin-top:54px;font:11px "DM Mono",monospace;color:#87918b;text-transform:uppercase;letter-spacing:.1em}
.hero-meta span:first-child{color:var(--accent)}
.hero-image{height:600px;margin:0;position:relative;background:#161d1a;border:1px solid var(--line)}
.hero-image figcaption{position:absolute;bottom:0;left:0;right:0;padding:15px 17px;display:flex;justify-content:space-between;background:linear-gradient(transparent,rgba(0,0,0,.85));font:10px "DM Mono",monospace;text-transform:uppercase;letter-spacing:.08em}
.section{padding:120px 0;border-bottom:1px solid var(--line)}
.section-label{margin-bottom:38px;color:#7f8983}
.split-heading,.section-intro{display:grid;grid-template-columns:1.15fr .85fr;gap:72px;align-items:start}
h2{font-size:clamp(42px,5.5vw,70px);line-height:.98;letter-spacing:-.055em;font-weight:500}
.split-heading p,.section-intro p{font-size:17px;color:var(--muted);max-width:520px}
.approach-text{max-width:780px;color:#d2d7d3;font-size:18px;margin:42px 0}
.pipeline{display:grid;grid-template-columns:1fr auto 1fr auto 1fr auto 1fr;align-items:center;border-top:1px solid var(--line);border-bottom:1px solid var(--line)}
.pipeline span{padding:22px 8px;font:11px "DM Mono",monospace;text-transform:uppercase;letter-spacing:.08em}
.pipeline i{font-style:normal;color:#69736d}
.project{margin-top:76px;padding-top:1px}
.project+.project{margin-top:120px;border-top:1px solid var(--line);padding-top:90px}
.project-main{display:grid;grid-template-columns:.95fr 1.05fr;gap:72px;align-items:center}
.project-copy h3{font-size:clamp(32px,4vw,50px);line-height:1.03;letter-spacing:-.045em;font-weight:500;margin:14px 0 20px}
.project-copy>p:not(.project-kicker){color:var(--muted);font-size:16px;max-width:610px}
.role{display:grid;grid-template-columns:70px 1fr;gap:18px;margin:28px 0;color:#b5beb9;font-size:12px}
.role span,.deliverables span,.about-columns span,.research-links>span{font:10px "DM Mono",monospace;color:#7d8881;text-transform:uppercase;letter-spacing:.1em}
.role strong{font-weight:500;color:#dce1dd}
.text-link{display:inline-flex;gap:10px;font-size:14px;font-weight:700;border-bottom:1px solid #5c665f;padding-bottom:5px}
.project-feature{height:520px;margin:0;background:#161c19;border:1px solid var(--line)}
.gallery{display:grid;grid-template-columns:repeat(6,1fr);gap:10px;margin-top:10px}
.gallery figure{margin:0;aspect-ratio:1/1;background:#121715;position:relative;overflow:hidden;border:1px solid var(--line)}
.gallery img{transition:transform .4s ease}
.gallery figure:hover img{transform:scale(1.04)}
.gallery figcaption{position:absolute;left:0;right:0;bottom:0;padding:26px 10px 9px;font:9px "DM Mono",monospace;text-transform:uppercase;letter-spacing:.08em;background:linear-gradient(transparent,rgba(0,0,0,.85))}
.service-grid{display:grid;grid-template-columns:repeat(3,1fr);margin-top:66px;border-top:1px solid var(--line);border-left:1px solid var(--line)}
.service-grid article{min-height:250px;padding:25px;border-right:1px solid var(--line);border-bottom:1px solid var(--line)}
.service-grid span,.steps>article>span,.expertise-list article>span{font:11px "DM Mono",monospace;color:var(--accent)}
.service-grid h3{font-size:22px;letter-spacing:-.025em;font-weight:600;margin:40px 0 12px}
.service-grid p,.steps p,.expertise-list p{font-size:14px;color:var(--muted)}
.deliverables{margin-top:28px;padding:20px 0;border-top:1px solid var(--line);display:grid;grid-template-columns:180px 1fr;gap:20px}
.deliverables p{margin:0;color:#d6ddd8;font-size:14px}
.steps{display:grid;grid-template-columns:repeat(5,1fr);margin-top:70px;border-top:1px solid var(--line)}
.steps article{padding:24px 18px 0;border-right:1px solid var(--line)}
.steps article:first-child{border-left:1px solid var(--line)}
.steps h3{font-size:18px;margin:38px 0 12px}
.expertise-grid{display:grid;grid-template-columns:.8fr 1.2fr;gap:90px}
.expertise-list{border-top:1px solid var(--line)}
.expertise-list article{display:grid;grid-template-columns:48px 1fr;gap:18px;padding:26px 0;border-bottom:1px solid var(--line)}
.expertise-list h3{margin:0 0 5px;font-size:18px}
.expertise-list p{margin:0}
.about{background:#0e1210}
.about-grid{display:grid;grid-template-columns:.65fr 1.35fr;gap:90px;align-items:start}
.about-image{height:560px;border:1px solid var(--line);background:#161c19}
.nameplate{margin:0 0 12px}
.about-copy h2{margin-bottom:28px}
.about-copy>p:not(.nameplate){max-width:720px;color:var(--muted)}
.about-columns{display:grid;grid-template-columns:1fr 1fr;gap:32px;margin-top:42px}
.about-columns div{border-top:1px solid var(--line);padding-top:16px}
.about-columns strong{display:block;margin-top:12px;font-size:13px;line-height:1.7;font-weight:500;color:#d8ded9}
.research-links{display:flex;flex-direction:column;gap:13px;margin-top:42px}
.research-links a{font-size:14px;color:#e2e7e3}
.research-links a:hover{color:var(--accent)}
.contact{padding-bottom:80px}
.contact-card{display:grid;grid-template-columns:1fr 1fr;gap:90px;background:#151b18;border:1px solid var(--line);padding:62px}
.contact-card h2{margin-bottom:24px}
.contact-card p{color:var(--muted);max-width:540px;font-size:15px}
.contact-email{display:inline-flex;margin-top:20px;font-size:18px;font-weight:700;border-bottom:1px solid #68736c;padding-bottom:5px}
.project-form{display:grid;grid-template-columns:1fr 1fr;gap:18px}
.project-form label{font-size:11px;color:#89938d;font:10px "DM Mono",monospace;text-transform:uppercase;letter-spacing:.08em}
.project-form label:nth-of-type(5),.project-form label:nth-of-type(6),.project-form button,.project-form small{grid-column:1/-1}
.project-form input,.project-form textarea{display:block;width:100%;margin-top:8px;background:#0c100f;border:1px solid #303934;color:#fff;padding:12px;border-radius:0;font:14px Manrope,Arial,sans-serif;outline:none}
.project-form textarea{resize:vertical}
.project-form input:focus,.project-form textarea:focus{border-color:#758078}
.project-form small{color:#7f8983;font-size:11px;line-height:1.5}
.project-form button{justify-self:start}
.site-footer{padding:34px 0 44px}
.footer-grid{display:grid;grid-template-columns:1fr auto 1fr;align-items:end;gap:30px}
.footer-grid>div:first-child{display:flex;flex-direction:column}
.footer-grid strong{font-size:14px}
.footer-grid span{font-size:11px;color:#7f8983;margin-top:4px}
.footer-grid nav{justify-content:center}
.footer-right{display:flex;justify-content:flex-end;gap:18px;font-size:11px;color:#89938d}
.footer-right a:hover{color:#fff}
@media(max-width:900px){
  .hero-grid,.project-main,.split-heading,.section-intro,.expertise-grid,.about-grid,.contact-card{grid-template-columns:1fr}
  .hero-image{height:480px}
  .project-feature{height:440px}
  .gallery{grid-template-columns:repeat(3,1fr)}
  .service-grid{grid-template-columns:1fr 1fr}
  .steps{grid-template-columns:1fr 1fr}
  .steps article:nth-child(3){border-left:1px solid var(--line)}
  .footer-grid{grid-template-columns:1fr}
  .footer-grid nav,.footer-right{justify-content:flex-start}
}
@media(max-width:620px){
  .container{width:min(var(--max),calc(100% - 30px))}
  .site-header nav a:not(.nav-cta){display:none}
  .hero{padding:62px 0 78px}
  .section{padding:80px 0}
  h1{font-size:54px}
  h2{font-size:43px}
  .hero-image{height:360px}
  .project-feature{height:340px}
  .gallery{grid-template-columns:repeat(2,1fr)}
  .steps article,.steps article:nth-child(3){border-left:1px solid var(--line)}
  .deliverables{grid-template-columns:1fr}
  .contact-card{padding:34px 24px;gap:52px}
  .footer-grid nav{gap:16px}
  .footer-right{flex-wrap:wrap}
}
.collaborations p{margin:13px 0 0;color:#d6ddd8;font-size:14px;max-width:720px}
.collaborations strong{font-weight:600;color:#e8ede9}

.institution-section{margin-top:42px;border-top:1px solid var(--line);padding-top:16px}
.institution-section>span{font:10px "DM Mono",monospace;color:#7d8881;text-transform:uppercase;letter-spacing:.1em}
.institution-grid{display:grid;gap:10px;margin-top:18px}
.collaboration-grid{grid-template-columns:repeat(3,1fr)}
.institution-chip{min-height:86px;border:1px solid var(--line);background:#111614;display:flex;align-items:center;gap:13px;padding:12px}
.institution-chip strong{font-size:11px;line-height:1.35;font-weight:600;color:#d8ded9}
.institution-mark{width:42px;height:42px;flex:0 0 42px;display:grid;place-items:center;border:1px solid #3a443e;background:#0b0f0e;color:#e6ebe7;font:10px "DM Mono",monospace;letter-spacing:-.02em}
.institution-mark.imperial{font-weight:500}.institution-mark.kings{border-radius:50%}
.institution-mark.nhm{font-size:9px}.institution-mark.unige{font-size:9px}
.institution-mark.phd{color:var(--accent);border-color:#4b5838}
.institution-mark.senckenberg{border-radius:50%;font-size:15px}
.institution-mark.uppsala{font-weight:600}
.institution-mark.flinders{border-radius:50%;font-weight:600}
@media(max-width:900px){}
@media(max-width:620px){}

.institution-chip div{display:flex;flex-direction:column;gap:4px}
.institution-chip small{font:9px "DM Mono",monospace;color:#7f8983;text-transform:uppercase;letter-spacing:.08em}

.credential-row{margin-top:18px;padding:18px 0;border-bottom:1px solid var(--line)}
.credential-row:first-child{margin-top:18px}
.credential-title{display:flex;align-items:center;gap:14px;margin-bottom:12px}
.credential-title strong{display:block;font-size:14px;color:#e1e6e2}
.credential-title small{display:block;margin-top:4px;font:9px "DM Mono",monospace;color:#7f8983;text-transform:uppercase;letter-spacing:.06em}
.institution-logos{display:grid;grid-template-columns:repeat(3,1fr);gap:10px;margin-left:56px}
.credential-row:nth-child(2) .institution-logos{grid-template-columns:repeat(2,1fr)}
.institution-logos .institution-chip{min-height:70px}
@media(max-width:900px){
  .institution-logos,.credential-row:nth-child(2) .institution-logos{grid-template-columns:1fr 1fr}
}
@media(max-width:620px){
  .institution-logos,.credential-row:nth-child(2) .institution-logos{grid-template-columns:1fr;margin-left:0}
}

.institution-chip{justify-content:flex-start}
.institution-chip strong{font-size:12px;line-height:1.45}
.credential-title{padding-left:0}

.research-links small{display:block;margin-top:4px;font:9px "DM Mono",monospace;color:#7f8983;text-transform:uppercase;letter-spacing:.06em}
.research-links .publication{display:flex;flex-direction:column;gap:5px;padding-top:13px;margin-top:4px;border-top:1px solid var(--line);max-width:760px}
.research-links .publication strong{font-size:12px;font-weight:600;color:#dfe5e1}
.research-links .publication span{font-size:13px;line-height:1.55;color:#c2cbc5}
.research-links .publication small{margin-top:0}

/* Selected publications */
.publication-list{display:grid;gap:0}
.publication-list a{display:flex;align-items:baseline;justify-content:space-between;gap:2rem;padding:1.25rem 0;border-top:1px solid rgba(255,255,255,.12);text-decoration:none}
.publication-list a:last-child{border-bottom:1px solid rgba(255,255,255,.12)}
.publication-list span{font-size:clamp(1rem,1.6vw,1.25rem);line-height:1.35}
.publication-list small{font-family:"DM Mono",monospace;font-size:.72rem;letter-spacing:.04em;white-space:nowrap;opacity:.7}
@media (max-width:700px){.publication-list a{display:block}.publication-list small{display:block;margin-top:.45rem}}
@media (max-width:700px){
}
@media (max-width:700px){
}
@media (max-width:700px){
}
@media (max-width:700px){
}
@media (max-width:700px){
}
@media (max-width:700px){}

/* Academic Background — rebuilt from scratch */
.academic-background{
  display:grid;
  grid-template-columns:repeat(2,minmax(0,1fr));
  gap:1rem;
  margin-top:1.5rem;
}
.academic-entry{
  padding:1.5rem 1.6rem;
  border:1px solid rgba(255,255,255,.12);
  border-radius:8px;
  background:rgba(255,255,255,.025);
}
.academic-degree{
  display:block;
  margin-bottom:1rem;
  font-family:"DM Mono",monospace;
  font-size:.75rem;
  letter-spacing:.08em;
  text-transform:uppercase;
  opacity:.65;
}
.academic-entry h3{
  margin:0;
  font-size:1.25rem;
  font-weight:600;
  letter-spacing:-.01em;
}
.academic-entry p{
  margin:.4rem 0 0;
  font-size:.88rem;
  opacity:.75;
}
@media (max-width:700px){
  .academic-background{grid-template-columns:1fr}
}

/* About — rebuilt as one coherent editorial section */
.about-subsection{margin-top:42px}
.about-kicker{display:block;font:10px "DM Mono",monospace;color:#7d8881;text-transform:uppercase;letter-spacing:.1em;margin-bottom:18px}
.about-card-grid{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:10px}
.about-card{min-height:118px;border:1px solid var(--line);background:#111614;padding:18px;display:flex;flex-direction:column;justify-content:flex-start}
.about-card-label{font:10px "DM Mono",monospace;color:#7d8881;text-transform:uppercase;letter-spacing:.1em;margin-bottom:18px}
.about-card h3{margin:0;font-size:14px;line-height:1.45;font-weight:600;color:#dfe5e1}
.about-card p{margin:7px 0 0;font-size:12px;line-height:1.5;color:#aeb8b1}
.collaboration-cards{grid-template-columns:repeat(3,minmax(0,1fr))}
.collaboration-cards .about-card{min-height:86px;justify-content:center}
.about-intro{max-width:720px;color:var(--muted)}
.about .research-links{margin-top:42px}
.about .research-links>span{font:10px "DM Mono",monospace;color:#7d8881;text-transform:uppercase;letter-spacing:.1em}
.about .publication-list{margin-top:18px}
.about .publication-list a{display:flex;align-items:baseline;justify-content:space-between;gap:24px;padding:15px 0;border-top:1px solid var(--line);text-decoration:none}
.about .publication-list a:last-child{border-bottom:1px solid var(--line)}
.about .publication-list a>span{font-size:14px;line-height:1.4;color:#e2e7e3}
.about .publication-list small{font:9px "DM Mono",monospace;color:#7f8983;text-transform:uppercase;letter-spacing:.06em;white-space:nowrap}
@media (max-width:900px){
  .collaboration-cards{grid-template-columns:1fr}
}
@media (max-width:700px){
  .about-card-grid{grid-template-columns:1fr}
  .about .publication-list a{display:block}
  .about .publication-list small{display:block;margin-top:5px}
}

.about-copy .collaboration-cards{grid-template-columns:repeat(2,minmax(0,1fr))}



/* Additional Works — preserve original narrow layout after relocation */
.work .research-links{
  width:100%;
  max-width:720px;
  margin-left:auto;
  margin-right:auto;
}

/* Section 02 — Selected Works */
.work .project-feature-wrap{display:block;width:100%}
.work .project-feature-wrap .project-feature{
  position:relative;
  margin:0;
  height:520px;
  overflow:hidden;
}
.work .project-feature-wrap .project-feature img{
  display:block;
  width:100%;
  height:100%;
  object-fit:cover;
}
.work .project-feature-caption,
.work .project-feature figcaption{
  position:absolute;
  left:0;
  right:0;
  bottom:0;
  z-index:3;
  display:block;
  margin:0;
  padding:34px 16px 12px;
  color:#fff;
  font:10px "DM Mono",monospace;
  line-height:1.5;
  letter-spacing:.06em;
  text-transform:uppercase;
  background:linear-gradient(to bottom,transparent,rgba(0,0,0,.88));
}
.work .zoomable{cursor:zoom-in}
.work .gallery{grid-template-columns:repeat(5,1fr);gap:10px}
.work .gallery figure{aspect-ratio:1/1}
.work .gallery img{width:100%;height:100%;object-fit:cover;display:block}
.work .gallery figcaption{z-index:2}
@media (max-width:1100px){
  .work .gallery{grid-template-columns:repeat(4,1fr)}
}
@media (max-width:900px){
  .work .project-feature-wrap .project-feature{height:440px}
  .work .gallery{grid-template-columns:repeat(3,1fr)}
}
@media (max-width:600px){
  .work .project-feature-wrap .project-feature{height:340px}
  .work .gallery{grid-template-columns:repeat(2,1fr)}
}


/* Image lightbox */
body.lightbox-open{overflow:hidden}
.image-lightbox{
  position:fixed;
  inset:0;
  z-index:9999;
  display:flex;
  align-items:center;
  justify-content:center;
  padding:clamp(18px,4vw,48px);
  background:rgba(5,7,8,.94);
  opacity:0;
  visibility:hidden;
  pointer-events:none;
  transition:opacity .2s ease,visibility .2s ease;
}
.image-lightbox.is-open{
  opacity:1;
  visibility:visible;
  pointer-events:auto;
}
.image-lightbox .lightbox-inner{
  width:min(94vw,1400px);
  height:min(90vh,900px);
  display:flex;
  flex-direction:column;
  align-items:center;
  justify-content:center;
  gap:14px;
}
.image-lightbox img{
  display:block;
  width:auto;
  height:auto;
  max-width:100%;
  max-height:calc(90vh - 54px);
  object-fit:contain;
  cursor:zoom-out;
  box-shadow:0 20px 70px rgba(0,0,0,.45);
}
.image-lightbox .lightbox-caption{
  width:100%;
  text-align:center;
  color:var(--muted);
  font:11px "DM Mono",monospace;
  line-height:1.5;
  letter-spacing:.06em;
  text-transform:uppercase;
}
.lightbox-close{
  position:absolute;
  top:18px;
  right:22px;
  z-index:2;
  width:44px;
  height:44px;
  border:1px solid rgba(255,255,255,.2);
  border-radius:50%;
  background:rgba(255,255,255,.06);
  color:#fff;
  font:32px/40px Arial,sans-serif;
  cursor:pointer;
}
.lightbox-close:hover{background:rgba(255,255,255,.12)}
.lightbox-close:focus-visible{
  outline:2px solid var(--accent);
  outline-offset:3px;
}
.work .zoomable img{
  cursor:zoom-in;
}
@media (max-width:600px){
  .image-lightbox{padding:18px}
  .image-lightbox .lightbox-inner{width:96vw;height:88vh}
  .image-lightbox img{max-height:calc(88vh - 50px)}
  .lightbox-close{top:10px;right:10px}
}

/* Additional Works — closing publication list for Selected Work */
.work .additional-works{
  width:min(100%, 760px);
  margin:clamp(64px,8vw,104px) auto 0;
  padding-top:clamp(28px,4vw,44px);
  border-top:1px solid var(--line);
}
.work .additional-works-heading{
  margin-bottom:24px;
}
.work .additional-works-heading .section-kicker{
  display:block;
  margin-bottom:8px;
  color:var(--accent);
  font:10px "DM Mono",monospace;
  letter-spacing:.12em;
  text-transform:uppercase;
}
.work .additional-works-heading h3{
  margin:0;
  font:600 clamp(22px,2.5vw,30px)/1.15 "Manrope",sans-serif;
  letter-spacing:-.025em;
}
.work .additional-works .publication-list{
  width:100%;
  max-width:none;
}
.work .additional-works .publication-list a{
  display:grid;
  grid-template-columns:minmax(0,1fr) auto;
  gap:18px 28px;
  align-items:start;
  padding:18px 0;
  border-top:1px solid var(--line);
}
.work .additional-works .publication-list a:last-child{
  border-bottom:1px solid var(--line);
}
.work .additional-works .publication-list a span:first-child{
  min-width:0;
}
.work .additional-works .publication-list a span:last-child{
  white-space:nowrap;
}
@media (max-width:700px){
  .work .additional-works{
    width:100%;
    margin-top:52px;
  }
  .work .additional-works .publication-list a{
    grid-template-columns:1fr;
    gap:8px;
  }
}

/* Video items in Selected Work use the same visual tile/caption treatment as images. */
.work .media-video{
  position:relative;
  margin:0;
  overflow:hidden;
}
.work .media-video video{
  display:block;
  width:100%;
  height:auto;
  aspect-ratio:4 / 3;
  object-fit:cover;
  cursor:zoom-in;
  background:#080a0b;
}
.work .media-video figcaption{
  display:block;
  margin-top:10px;
  color:var(--muted);
  font:10px "DM Mono",monospace;
  line-height:1.5;
  letter-spacing:.06em;
  text-transform:uppercase;
}
.image-lightbox .lightbox-video{
  display:block;
  width:auto;
  height:auto;
  max-width:100%;
  max-height:calc(90vh - 54px);
  object-fit:contain;
  cursor:default;
  box-shadow:0 20px 70px rgba(0,0,0,.45);
}

/* Small top edge shadow for the two large Selected Work feature images,
   matching the compact bottom caption shadow treatment. */
.work .project-feature::before{
  content:"";
  position:absolute;
  left:0;
  right:0;
  top:0;
  height:8%;
  pointer-events:none;
  z-index:2;
  background:linear-gradient(to bottom,rgba(0,0,0,.85),transparent);
}